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    The Hardness locus on chromosome 5D is the main determinant of grain texture in hexaploid wheat. The related genes Puroindoline-a and -b and Grain Softness Protein are tightly linked at this locus. Mutations in the Pina-D1 and Pinb-D1 genes are associated with increased grain hardness. We report here the complete sequence of a 101-kb BAC clone from Triticum monococcum which includes these three genes, and its comparison with the orthologous region in rice. The genes Gsp-A m1, Pina-A m1 and Pinb-…Read more